- 1 The domestic life of cats (history)
- 2 The features of the cat
- 3 Making a home for a cat
- 4 Colonies
- 5 Cats in the wild, and feral cats
- 6 Overpopulation
- 7 Varieties of domestic cat
- 8 Coat patterns
- 9 References
| Felis catus|
Litter mates, or even generations of house cats with a common parent, may be closely bonded. Kittens handled lovingly in their first weeks are apt to bond with humans. They tend to form their food preferences in their first six months, and most cats are picky eaters — although some develop fascinating tastes, from melon to cream cheese to potatoes.
While cats are often considered aloof, individual cats and individual humans form intense human-animal bonds.
Adult males (called tomcats or toms) operate independently, associating only to fight other males and court and mate with females. Though the house cat has changed over the thousands of generations the species has lived with and around us, and the modern cat is more inclined to associate with other adults of its kind than its wild ancestors (see the section on cat colonies below), the cat has never been, and is not now, inclined to accept orders from any kind of group leader. Cats only pay homage to the head of the household perhaps in the form of headless mice deposited at the front door, and not because a leader asked for them. Indeed, this behaviour may stem less from tribute than from an instinctive inclination to share the remains of the hunt with its less capable family members, or to teach the "kittens" what they should be doing.
The technical word for a group of cats, "clowder," is used, but typically among specialists and for large groups.
The domestic life of cats (history)
Why did we adopt such a singular creature into our homes and families?
Cats' skill at exterminating rodents practically benefits humans the most. The domestic cat, unlike most predators (including the big cats), does not hunt and kill only when hungry. Instead, the typical house cat follows the lure of the chase for its own sake. Most follow a scampering mouse or fluttering bird with focused abandon, even if completely sated. To see a cat stalk a bird, the tip of its tail twitching, the mouth spasmodically gaping, is to witness a form of intoxication. This addiction to the hunt, along with sharp teeth, an athletic pounce, silent footfalls, and sharp claws combine to make the cat a relentless assassin, who, even if gorged on prey, will continue with its work. The apparent cruelty of cats has caused many people to disdain them, but their obsession with stalking and "playing" with little moving things probably led to their domestication in the first place.
Historians document cats' first domestic presence in ancient Egypt, about 1500 BC, where granaries first appeared. Mice and other rodents multiply around large stores of grain, making cats desirable for ancient Egyptians, which evidence indicates. The ancient Egyptians mummified cats as lavishly as royalty and aristocrats, and records indicate they considered killing or harming a cat a serious crime.
The cat is one of the few domestic animals that adapts easily to shipboard life; that ability, along with a talent for the extermination of vermin, has accounted for the long term global expansion of this animal's territory. In other words, cats that managed to board seagoing vessels—invited or otherwise—have been tolerated and have thrived, seeding cat populations throughout the world even when not deliberately imported.
The features of the cat
Cats thrive in most homes.
Weighing on average between 2.5 and 7 kg (5.5–16 pounds), the narrow range of size between the vast majority of healthy weight adult cats differs from the great size differences between the largest and smallest breeds of dogs or horses. Purebred cats, a small minority, tend to represent the extremes of size and other features. The Maine Coon , for example, is one of the largest and heaviest, sometimes 11.3 kg (25 pounds) in lean weight and the Korat  one of the smallest, only about 3 kg (6-7 lbs).
Cats live 10 to 20 years, and usually longer if owners keep them indoors. Restricting the cat to an indoor life reduces the risk of injury from other cats, dogs and wild predators, as well as motor vehicle accidents and exposure to diseases. Spayed or neutered cats also tend to live longer. In captivity, well-cared for indoor cats often live 14 to 20 years, with 36 years being the record for the oldest pet cat. Although cats given the freedom to go outside tend to die younger, some of these inside/outside cats live a long time.
Eyes and vision
Testing indicates cats see better at night than humans, but not as well in daylight. The cat's tapetum lucidum, as with dogs and many other animals, reflects extra light to the retina. A camera's flash interacts with the tapetum to make the cat's eye color vary in photographs. While this structure enhances the ability to see in low light, it reduces net visual acuity, and so the cat's pupil can close enough to eliminate much of the incoming daylight. In very bright light, the slit-like iris closes very narrowly over the eye, reducing the light on the sensitive retina, and improving depth of field. Cats can thus detect light seven times more sensitively than humans.
Cats' field of view, estimated at 200° (20° wider than humans'), has a narrower central binocular field (the overlap in the images from each eye) than humans'. As with most predators, including us, their eyes face forward, affording depth perception at the expense of field of view, which largely depends upon the eyes' placement, but may also relate to the eye's construction. Instead of the fovea which gives humans sharp central vision, cats have a central band known as the visual streak. Cats can apparently differentiate among colors, especially at close range, but without appreciable subtlety. Like many animals, the cat's nictitating membrane, a translucent "third eyelid," closes from the side to thinly cover the eye, and appears when the cat's eyelid first opens. A sick cat's nictitating membrane partially closes, but a sleepy, content cat's membrane is often visible. If a cat chronically shows the third eyelid, it should be taken to a veterinary surgeon.
Ears and hearing
Humans and cats hear low frequencies similarly, but cats can hear much higher-pitched sounds - up to 64,000 kHz, which is 1.6 octaves above the range of a human, and even an entire octave above the range of a dog. In most animals, the ear is "tuned" to hear the sounds most important for species survival, and that means the sounds of its kind, its prey and its predators. Cats' excellence, for example, at hearing faint, high-pitched sounds helps them locate rodents' nests. Cats swivel their ears toward the direction they're listening, and a cat's ear flaps (pinnae) can independently point backwards, forwards, and sideways to pinpoint the source of the sound. Cats can judge within three inches (7.5 cm) the location of a sound made one yard (approximately one meter) away - another attribute adding to their consummate skill at hunting.
Sense of smell
Cats have twice as many smell-sensitive cells in their noses as people, and can likely smell odors we never sense. The cat's scent organ in the roof of its mouth, called the vomeronasal, or Jacobson's organ, causes extreme sensitivity toward open flames, fire, or burning objects. When a cat wrinkles its muzzle, lowers its chin, and lets its tongue hang a bit, it is opening the passage to the vomeronasal. This is called gaping, "sneering", or "flehming", which equates to the Flehmen response in other animals, such as dogs, horses and big cats.
Cats generally have about a dozen whiskers in four rows on each upper lip, a few on each cheek, tufts over the eyes, and bristles on the chin. Whiskers may also be on the cat's inner "wrists", and similar hairs make up the cat's eyebrows. The Sphynx (a nearly hairless breed) may have full length, short, or no whiskers at all.
Whiskers (called vibrissae) can aid with navigation in the dark. Whiskers may detect very small shifts in air currents, alerting a cat to an unseen obstruction's proximity. The upper two rows of whiskers can move independently from the lower two rows for even more precise measuring. These whiskers also spread out roughly as wide as the cat's body, enabling it to judge if it can fit through an opening. Cats also have a number of reinforced hairs similar to whiskers on other parts of their bodies: over the eyes, on the chin and at the back of the legs. 
Cats, like dogs, are digitigrades: they walk directly on their toes, the bones of their feet making up the lower part of the visible leg. Cats can walk very precisely because like all felines they directly register, that is, they place each hind paw almost directly in the print of the corresponding forepaw. Such walking minimizes noise and visible tracks and provides sure footing for their hind paws when navigating rough terrain.
Unlike dogs, cats walk by moving both legs on one side and then both legs on the other, a type of 'pacing gait'. This is their normal walk; they have an extensive range of jumps and twists.
Like all members of family felidae, with the exception of the cheetah, house cats have retractable claws. In the normal, relaxed position the skin and fur around the toe pads sheaths the claws, keeping them sharp by preventing wear from contact with the ground and allowing the silent stalking of prey. The forefoot's claws are usually sharper than the hindfoot's. Cats can extend their claws voluntarily on one or more paws at will. Cats may extend their claws in hunting or self-defense, climbing, "kneading", or for extra traction on soft surfaces (bedspreads, thick rugs, etc.).
Most cats have four claws on each of their hind paws and five on their front paws, but an allele in the gene pool makes them prone to polydactyly, and some have six or seven toes. One breed, the Maine Coon, has a particularly high rate of polydactyly.
Coat and eye color
Cats come in a variety of colors, solid colored or striped.
Cats not only have many different eye colors, most typically yellow, copper, green, and orange, but some (odd eyed) varieties even feature two different colors. Purebred cats' eye color is often specified as a feature of the breed standard. Blue eyes are associated with the Siamese breed, though white cats also have them. White cats with two blue eyes are often deaf; orange eyes, however, usually indicate the cat hears normally. White cats having one blue and one other-colored eye are called "odd-eyed" and may be deaf on the same side as the blue eye.
Body language of cats
Normally born in a litter and raised by the mother, cats communicate with each other vocally and bodily. Cats also emit odors other cats interpret, but humans can't consciously understand these signals. The facial expressions, sounds, body postures and movements of paw and tail, however, can be read.
Whiskers also indicate the cat's attitude, pointing forward when the cat is inquisitive and friendly, and lying flat back on the face when defensive or aggressive. Whereas a dog wags its tail to signal friendly intentions, the motions of a cat's tail are more complex. Twitching the tip may mean curiosity or a greeting, or, snapped rapidly, hunting reflex. Wide sweeps of the entire tail show anger.
Normally between 38 and 39 °C (101 and 102.2 °F), a cat's body temperature is considered febrile (hyperthermic) when above 39.5 °C (103 °F), or hypothermic if below 37.5 °C (100 °F). For comparison, humans have a normal temperature of approximately 36.8 °C (98.2 °F). A domestic cat's normal heart rate ranges from 140 to 220 beats per minute, and largely depends on how excited the cat is. For a cat at rest, the average heart rate should be between 150 and 180 bpm, about twice that of a human.
Making a home for a cat
Most breeds of cat like to settle in high places, or perch.
The litter box
Owners may provide indoor cats a litter box containing sand or similar material (litter) which cats use the way humans use a toilet. Owners should clean the box daily and change the litter often, depending on the number of cats in a household and the type of litter; clumping litter stays cleaner longer, but reportedly causes health problems in some cats.  A litter box is recommended for indoor-outdoor cats as well. Litter boxes may risk transmitting toxoplasmosis to susceptible pregnant women and immuno-compromised individuals, but cleaning the litter box daily reduces the risk when cleaned by someone other than child-bearing women.
Hunting and diet
Even the most well-fed domestic cats will hunt and kill birds, mice, rats and other small animals when given the opportunity. They often present such trophies to their owner. The motivation is not entirely clear, but friendly bonding behaviors are often associated with such an action. Their ceaseless hunting makes cats hazardous to small animals, including locally endangered bird species. In some cases, cats have contributed to or caused extinctions -— for example, see the case of the Stephens Island Wren. So it is not just the cat who lives longer when it is kept indoors and not allowed free access outside.
Still, even an indoor cat has access to prey in many homes. Although some individual house cats have tolerated the company of birds (even allowing close contact with small birds), generally any small pet must be kept away from the cat. Cats can terrorize small creatures in cages, even when they cannot actually physically reach them. Wild rodents inside the home are, of course, fair game to a house cat and many owners keep cats for their skill at eliminating vermin indoors. Whether an owner approves or disapproves, however, this activity is likely to go on as long as the cat shares a home with mice and their kind. The cat having contact with wild rodents and other wild animals making their way into the home (like bats) is potentially exposed to certain parasites (for example, tapeworms), and both bacteria and viruses that can cause disease, so even strictly indoor cats require vaccination from such diseases as rabies, and cleaning out the cat box may spread toxoplasmosis to a human owner.
In captivity, cats cannot live on an unsupplemented vegetarian diet because they cannot synthesize several needed nutrients absent or rare in plant food, mainly taurine, vitamin A (cats cannot convert the pro-vitamin A that is abundant in plants to vitamin A proper) and certain fatty acids. The absence of taurine causes the cat's retina to slowly degenerate, causing eye problems and (eventually) irreversible blindness. This condition is called central retinal degeneration (CRD). Cow's milk is a poor source of taurine and adult cats are generally lactose intolerant. Lactose-free milk is perfectly safe, but still doesn't substitute for meat.
In nature, cats are almost never overweight, but obesity is recognized as a health problem among pet cats.
Some houseplants can harm cats. The leaves of the Easter Lily can cause permanent and life-threatening kidney damage. Philodendron are also poisonous to cats. Cat Fancy has a full list of plants harmful to cats.
Most cats are fond of catnip. While they generally do not consume it, they will often roll in it, paw at it, and occasionally chew on it (as catnip is sensed by the cat's Vomeronasal organ). The effect usually lasts only a few minutes. After two hours or less, susceptible cats gain interest again. Several other species of plants cause this effect, to a lesser degree.
Because of their small size and limited persistence in any attack, cats pose little direct danger to humans — the main hazard is the possibility of infection (e.g., cat scratch disease, or, rarely, rabies) from a cat bite or scratch.
Cats groom fastidiously by licking their fur and employing their hooked pappilae and saliva. Their saliva is a powerful cleaning agent, but it can provoke allergic reactions in humans. Many cats also enjoy grooming humans or other cats. Some cats occasionally regurgitate hair balls of fur that grooming collects in their stomachs, to which longhair cats are more prone to than shorthairs. Hairballs can be prevented with certain cat foods and remedies that ease eliminating the hair and by grooming the coat regularly with a comb or stiff brush. Cats expend nearly as much fluid grooming as urinating.
Some cats have been successfully toilet trained, eliminating the litterbox and its attendant expense and smell.
Indoor cats provided a scratching post are less likely to ruin furniture with their claws. No matter how much some cats may use a scratching post, other cats do not use them at all.
Declawing, a major surgery known as onychectomy performed under anesthesia, removes the tip of each digit (from the first knuckle out) of the cat's forepaws (and sometimes the hind paws). Veterinarians generally criticize the procedure and some refuse to perform it. Declawing surgery can be done by incision or laser. Laser declawing is considered more humane because although done under anesthesia, the cat normally recovers much faster because it bleeds minimally and the laser both removes the claw and cauterizes the wound.
The Cat Genome Project, sponsored by the Laboratory of Genomic Diversity at the U.S. National Cancer Institute Frederick Cancer Research and Development Center in Frederick, Maryland, focuses on the cat as an animal model for human hereditary disease, infectious disease, genome evolution, comparative research initiatives within the family Felidae, and forensic potential.
WalkingUnlike dogs, who show deference and are obedient, regarding their human as the leader of the pack, cats are independent-minded and prefer to select where to go and what to study and explore. It's usually easy to take a dog for a walk, even without a lead, but difficult with a cat. However, according to one source, it is possible to teach some cats to walk on a leash by using this method:
The trick is (usually) starting early, and then using a small dog harness that spreads the pull, rather than a collar.
Despite its reputation as a solitary animal, the domestic cat socializes enough to form colonies, but does not attack in groups as do lions. Some breeds like bengal, ocicat and manx are very social, but these are exceptions. While each cat holds a distinct territory (sexually active males having the largest territories, and neutered cats having the smallest), cats respect "neutral" areas where they watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually aggressively chase away stranger cats, at first by staring, hissing, and growling, and if that does not work, by short but noisy and violent attacks. Fighting cats raise their fur and arch their backs to increase their visual size and thus make themselves appear more impressive and threatening. Cats also do this while playing. Attacks usually comprise powerful slaps to the face and body with the forepaws as well as bites, but rarely damage seriously; usually the loser runs away with little more than a few scratches to the face, perhaps the ears. Normally, serious negative effects are limited to possible infections of the scratches and bites, though these sometimes kill cats if not treated. Sexually active males will usually be in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to the ears and nose. Not only males will fight; females will also fight over territory or to defend their kittens, and even neutered cats will defend their (smaller) territories aggressively.
Cats in the wild, and feral cats
Unlike the wild counterpart to other common domesticated animals, like wild horses and wolves, the original undomesticated wild cat has continued to flourish in the wild. Meanwhile, the population of domesticated cats has skyrocketed. Not all domestic cats are cared for as pets, some living in barns carrying on the tradition of the first domesticated cats in the Egyptian graineries. More, however, are strays; abandoned, they often survive long enough to breed and, unlike their wild counterparts, commonly remain together in groups as the kittens grow. Stray and abandoned pet cats are a world-wide problem. In 1995, local municipalities throughout Japan put down at least 307,626 cats.
Feral cats may live alone, but most live in large groups called feral colonies with communal nurseries, depending on resource availability. Urban areas in the developed world are not friendly, nor adapted, environments for cats; most domestic cats descend from cats in desert climates and were distributed throughout the world by humans. Nevertheless, some feral cat colonies are found in large cities, e.g., around the Colosseum and Forum Romanum in Rome.
However, thousands of volunteers and organizations trap these unadoptable feral felines, spay or neuter them, immunize the cats against rabies and feline leukemia, and treat them with long-lasting flea products. Before releasing them back into their feral colonies, the attending veterinarian often nips the tip off one ear to mark the feral as spayed/neutered and inoculated, as these cats will likely get trapped again. Volunteers continue to feed and care for these cats throughout their lives, greatly increasing their lifespan and reducing behavior and nuisance problems due to competition for food. In time, if an entire colony is successfully spayed and neutered, no additional kittens are born and the feral colony disappears. Many hope these efforts will end urban feral cat colonies.
According to the Humane Society of the United States, 3-4 million cats and dogs are euthanized each year in the United States and many more confined to cages in shelters because significantly more animals are born than there are homes. Spaying or neutering pets helps reduce overpopulation.  Local humane societies, SPCA's and other animal protection organizations urge people to spay or neuter their pets and to adopt animals from shelters instead of purchasing them.
Varieties of domestic cat
Cat breeds have distinct features and heritage. Most cat registries recognize between 35 and 70 breeds, and several more are in development, with one or more new breeds recognized each year on average. When choosing a cat, a purebred or mixed breed cat should be considered. The owners and breeders of show cats compete to see whose animal most closely resembles the "ideal" definition of the breed (see selective breeding). Due to common crossbreeding in populated areas, many cats are simply identified with the homogeneous breeds of domestic longhair and domestic shorthair, depending on their fur. British and Australian slang refer to non-purebred cats as moggies, which derives from "Maggie", short for Margaret, reputed to have been a common name for cows and calves in 18th-century England and later applied to housecats during the Victorian era. In the United States, a non-purebred cat is sometimes referred to in slang as a barn or alley cat, even if it is not a stray.
Cats come in a variety of colors and patterns. These are physical properties and should not be confused with a breed of cat.
Household cats are divided into:
Cat coat genetics can produce a variety of coat patterns. Some of the most common are:
- Bicolor, Tuxedo and Van
- This pattern varies between the tuxedo cat, which is mostly black with a white chest, and possibly markings on the face and paws/legs, all the way to the Van pattern (named after the Lake Van area in Turkey, which gave rise to the Turkish Van breed), where the only colored parts of the cat are the tail (usually including the base of the tail proper), and the top of the head (often including the ears). There are several other terms for amounts of white between these two extremes, such as harlequin or jellicle cat. Bicolor cats' primary (non-white) color can be black, red, or any dilution thereof, or tortoiseshell (see below for definition).
- Tabby cat
- Striped, with a variety of patterns. The classic "blotched" tabby (or "marbled") pattern is the most common and consists of butterflies and bullseyes. The "mackerel" or "striped" tabby pattern is a series of vertical stripes down the cat's side (resembling the fish). This pattern broken into spots is a "spotted" tabby. Finally, the tabby markings may look like a series of ticks on the fur, hence the "ticked" tabby, which is almost exclusively associated with the Abyssinian breed. The worldwide evolution of the cat means that certain types of tabby are associated with certain countries; for instance, blotched tabbies are quite rare outside NW Europe, where they are the most common type.
- Feline Statistics. Retrieved on August 15, 2005.
- Strain, G.M., How Well Do Dogs and Other Animals Hear?
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- Normal Values For Dog and Cat Temperature, Blood Tests, Urine and other information in ThePetCenter.com. Retrieved on August 8, 2005.
- Suspected bentonite toxicosis in a cat from ingestion of clay cat litter. Retrieved on September 10, 2005.
- The plight of abandoned pets: Thrown out with the city's garbage Kyoko Sato. Japan Times (International). Tokyo: Feb 9-Feb 15, 1998. Vol. 38, Iss. 6; p.7
- "Moggie" definition question and answer Worldwidewords.org. URL Accessed June 14, 2006.